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Species Information
Summary: Features include a hoof-shaped to spherical, tan-colored fruitbody on conifers, which when sliced open is seen to have a hollow cavity lined in its upper part by pores, a tube layer and flesh, and in its lower part by a relatively thin layer that may be perforated. Spores are apparently spread by bark beetles as well as by wind.
It is found in BC, WA, OR, ID, AB, MB, NB, ON, AZ, CA, CO, DE, FL, MA, MD, ME, MI, MN, MT, NE, NH, NJ, NM, NV, NY, PA, SD, UT, VA, VT, WV, WY, and east Asia, (Gilbertson).
Cap: 1.5-8.5cm, more or less spherical to ovoid or slightly compressed or hoof-like, with hollow interior, upper surface whitish to warm tan or yellowish drying ocher-brown to reddish brown, with thin, smooth, glazed or resinous crust, ''margin extending down and under to form "veil" which completely covers pore surface; in age the underside perforated by one (rarely two) holes'', (Arora), up to 4cm x 5cm x 4cm, hoof-shaped, margin continuous with a volva-like structure that encloses pore surface except for a small hole at the base; upper surface cream to yellowish or tan; not zoned, bald but often coated with a clear lacquer-like layer, smooth or wrinkled, (Gilbertson)
Flesh: tough; whitish, (Arora), up to 2cm thick, soft-corky; ivory-white, not zoned, (Gilbertson)
Pores: 3-4 per mm, hidden by "veil", "white becoming pinkish or brownish in age", (Arora), 4-5 per mm, circular, with thick, entire walls; pale to dark chocolate brown; tube layer up to 0.6cm thick, pinkish buff, (Gilbertson)
Microscopic: spores 8-12 x 3-5 microns, cylindric to elliptic, smooth, (Arora), spores 12-16.5 x 4.5 microns, cylindric, smooth, inamyloid, colorless; cystidioles 20-28 x 5-7 microns, not projecting, but fusoid, thin-walled, with basal clamp; hyphal system trimitic, contextual generative hyphae mostly 3-7 microns wide (but with inflated parts at branches up to 15 microns wide), thin-walled, with clamp connections at all septa, with occasional branching, contextual skeletal hyphae 2.5-8 microns, thick-walled, colorless, nonseptate, with occasional branching, binding hyphae 1.5-2.5 microns wide, thick-walled, nonseptate, much branched, tramal hyphae similar, (Gilbertson)
Spore Deposit: pinkish or flesh-colored (Arora), white to cream-color (Miller)
Habitat / Range
annual, "solitary or more often in groups on dead or occasionally old living conifers", (Arora), single or in large numbers, on recently killed conifers, "commonly fruiting a year after trees are killed by fire, bark beetles, or other factors and then displaced by other saprophytic wood-rotting fungi", especially on Pinus (pine), Abies (fir), and Pseudotsuga (Douglas-fir), causes soft grayish-white rot of sapwood, (Gilbertson), fruiting in spring, summer, and fall, (Miller)
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